Xylemtissue that carries water up the root and stem. In trees, it constitutes wood — hence the word is derived from Greek ξύλον xúlon "wood, timber". Together with phloem, xylem is one of the two transport tissues of plants. The cell walls of xylem cellss derive most of their strength from lignin, a chemical compound produced only by plants.
Xylem (at least in dicots) is composed of vessel elementss and tracheids. Vessel elements are similar in structure to the sieve-tube members of phloem, but they lack companion cells, and have perforated sides as well as pores at the ends. Tracheids are much narrower cells, with tapered and perforated ends, constituting most of the volume of the xylem tissue. Both tracheids and vessel elements are dead at maturity.
- The soil solution (see soil) is more dilute than the cytosol of the root cells. Thus, water moves osmotically into the cells, creating root pressure. Even under optimal conditions, root pressure can only lift water a couple of feet.
- Capillary action helps sap to flow up the narrow tracheids.
- By far the most important cause of xylem sap flow is transpirational pull. This is the reverse of root pressure, caused by the transpiration of water from leaves.